Isn't the grinning one in the middle on the left one of the moderators on the old board? (All Australians look alike.)
Scientists from Georgia Tech are on the case.
It appears to be the result of how the lower intestines in these animals expand as the bowels are filled with solidifying proto-poop. Kind of a biological injection-molding, except that the mold is flexible tissue that expands to take its square shape as the turd-material is injected from the rest of the intestine into it.
https://phys.org/news/2018-11-scientists...-poop.html